- Leathery leaves.
- Pinnately compound.
- Pinnule (leafule) with triangular auricle at base.
- Fertile pinnae (leaflets) on upper part of frond (leaf) and are much smaller than lower, non-fertile pinnae.
- Sori covers fertile pinnae.
- Two lowest pinnae often down-pointed.
- Hairs/spines at tip of pinnule teeth.
